1 Departments of Medicine and Cell and Developmental Biology, Abramson Family Cancer Research Institute, Perelman School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania 19104, USA.
2 Experimental Medicine I, Nikolaus-Fiebiger-Center for Molecular Medicine, FAU University of Erlangen-Nuernberg, 91054 Erlangen, Germany.
3 Department of Molecular Biology, Princeton University, Princeton, New Jersey 08544, USA.
4 Instituto de Neurociencias CSIC-UMH, Avda. Ramón y Cajal s/n, 03550 San Juan de Alicante, Spain.
5 Whitehead Institute for Biomedical Research, Cambridge, Massachusetts 02142, USA.
6 Ludwig/MIT Center for Molecular Oncology, Cambridge, Massachusetts 02142, USA.
7 Department of Biology, Massachusetts Institute of Technology, Cambridge, Massachusetts 02142, USA.