1 College of Forestry, Henan Agricultural University, Zhengzhou, 450002, China.
2 Liaoning Key Laboratory for Biological Invasions and Global Change, Shenyang Agricultural University, Shenyang, Liaoning, 110866, China.
3 Guangdong Provincial Key Laboratory of Soil and Groundwater Pollution Control, School of Environmental Science and Engineering, Southern University of Science and Technology, Shenzhen, Guangdong, 518055, China.
4 International Center of Tropical Botany, Florida International University, Miami, FL, 33199, USA.
5 Department of Forest Ecology and Management, Swedish University of Agricultural Sciences, Umeå, 90183, Sweden.