1 Versus Arthritis Centre for Genetics and Genomics, Centre for Musculoskeletal Research, Manchester Academic Health Sciences Centre, The University of Manchester, Manchester, UK.
2 NIHR Manchester BRC, Manchester University Foundation Trust, Manchester, UK.
3 Versus Arthritis Centre for Epidemiology, Centre for Musculoskeletal Research, Manchester Academic Health Sciences Centre, The University of Manchester, Manchester, UK.
4 Leeds Institute of Rheumatic and Musculoskeletal Medicine, University of Leeds and NIHR Leeds Musculoskeletal Biomedical Research Unit, Leeds Teaching Hospitals NHS Trust, Leeds, UK.
5 Institute of Cellular Medicine, Newcastle University, Newcastle upon Tyne, UK.
6 National Institute for Health Research Newcastle Biomedical Research Centre at Newcastle upon Tyne Hospitals NHS Foundation Trust and Newcastle University, Newcastle upon Tyne, UK.
7 UCD School of Medicine and Medical Science, Conway Institute, University College Dublin, Dublin, Ireland.
8 Versus Arthritis Centre for Genetics and Genomics, Centre for Musculoskeletal Research, Manchester Academic Health Sciences Centre, The University of Manchester, Manchester, UK. darren.plant@manchester.ac.uk.
9 NIHR Manchester BRC, Manchester University Foundation Trust, Manchester, UK. darren.plant@manchester.ac.uk.