1 Center for Functional Anatomy and Evolution, Johns Hopkins University School of Medicine, 1830 E. Monument St., Baltimore, MD 21111, USA. Electronic address: cbruff@jhmi.edu.
2 Unité d'Anthropologie bio-culturelle, Droit, Éthique et Santé (ADÉS), UMR 7268, Université d'Aix-Marseille-EFS-CNRS, Marseille, France; Département de Préhistoire, MNHN, UMR 7194, Paris, France.
3 Département de Préhistoire, MNHN, UMR 7194, Paris, France; Département Géosciences, Université de Poitiers, France.
4 Department of Anatomy and Cell Biology, University of Iowa, Iowa City, IA 52242, USA.
5 Department of Anthropology and Museum of Natural History, University of Iowa, Iowa City, IA 52242, USA.