1 From the Wellcome Centre for Human Genetics, Nuffield Department of Medicine, University of Oxford, Roosevelt Drive Headington, Oxford, UK.
2 Oxford Vaccine Group, Department of Paediatrics, University of Oxford, Churchill Hospital Old Road Headington, Oxford, UK.
3 Chinese Academy of Medical Science (CAMS) Oxford Institute, Nuffield Department of Medicine, University of Oxford, Old Road Campus Headington, Oxford, UK.