1 School of Molecular & Cellular Biology, Faculty of Biological Sciences, University of Leeds, Leeds, UK; Radiotherapy Research Group, Leeds Institute of Medical Research at St James's, Faculty of Medicine & Health, University of Leeds, Leeds, UK; Leeds Cancer Centre, The Leeds Teaching Hospitals NHS Trust, Leeds, UK.
2 Department of Medical Physics & Biomedical Engineering, University College London, London, UK.
3 CRUK/MRC Oxford Institute for Radiation Oncology, University of Oxford, Oxford, UK.
4 Christie Hospital, The Christie NHS Foundation Trust, Manchester, UK.
5 Velindre Cancer Centre, Cardiff, UK. Electronic address: tom.crosby@wales.nhs.uk.